Env. Science
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AP Environmental Science — Cheatsheet

Formulas, exam-day tips, and key terms on one page.

Formulas & relationships

What separates biomes
terrestrial biome ≈ f(temperature, precipitation)
Aquatic biomes instead key on salinity, depth, and flow. Same idea: the abiotic setting selects the community.
The 10% rule
energy at next level ≈ 0.10 × energy at current level
A rule of thumb, not a law — real efficiencies range from ~5% to ~20%. AP problems assume exactly 10% unless told otherwise.
Key difference among the cycles
C and N: large atmospheric reservoir · P: no atmospheric phase
Carbon (CO₂) and nitrogen (N₂) both cycle through the air; phosphorus moves through rock, water, and soil only — which makes it move slowly.
Net primary productivity
NPP = GPP − R
R is the energy producers spend on their own respiration. NPP is what remains as biomass; GPP is always larger than NPP.
Productivity and trophic transfer
NPP = GPP − R energy at next level ≈ 0.10 × energy at current level
NPP is what consumers can eat. The 10% figure is an average; real efficiencies run from about 1% to 20%.
Simpson’s Diversity Index
D = 1 − [ Σ n(n − 1) / (N(N − 1)) ]
n = number of individuals of a given species; N = total individuals of all species. D ranges from 0 to nearly 1 — values closer to 1 mean higher diversity (more species and more even abundances).
Species equilibrium on an island
species richness balances immigration rate against extinction rate
Large islands lower extinction; near islands raise immigration. Both effects push species number up, so a large + near island is the most species-rich.
Tolerance and environmental change
wide tolerance (generalist) → survives change · narrow tolerance (specialist) → vulnerable
Rapid environmental change favors species with broad tolerance ranges. Specialists, tuned to narrow conditions, are the first to decline.
Primary vs. secondary succession
primary: starts with NO soil (bare rock) · secondary: starts with soil intact
The presence or absence of soil is the deciding test. Soil already present → secondary (fast). Bare rock, no soil → primary (slow).
Island biogeography equilibrium
species number equilibrates where immigration rate = extinction rate
Immigration rises with proximity to a source; extinction falls with island size. Large and near supports the most species.
Population growth rate
growth rate (%) = [(births + immigration) − (deaths + emigration)] / N × 100
N is the starting population. A positive result means growth; a negative result means decline. If migration is negligible, growth rate ≈ (CBR − CDR) / 10 as a percent.
Matching strategy to curve
r-selected → many offspring, little care → Type III · K-selected → few offspring, much care → Type I
r-strategists bet on quantity and rebound fast; K-strategists bet on quality and recover slowly. Type II (constant mortality) sits between the extremes.
Two kinds of limiting factors
density-dependent: competition, predation, disease · density-independent: weather, natural disasters
If the factor gets worse as the population gets more crowded, it is density-dependent. If it strikes regardless of crowding (usually abiotic events), it is density-independent.
Rule of 70 (doubling time)
doubling time (years) ≈ 70 / annual growth rate (%)
Enter the growth rate as a plain percent number (a 2% rate → 70/2 = 35 years). Only works for steady exponential growth, and the growth rate must be positive.
Growth rate and the rule of 70
growth rate = [(births + immigration) − (deaths + emigration)] / population doubling time ≈ 70 / (% growth per year)
Immigration and emigration are frequently omitted by students and are frequently in the data.
Replacement fertility
TFR ≈ 2.1 for replacement in a low-mortality country; higher where child mortality is high
Above 2.1 means eventual growth, below means eventual decline — but age structure delays both.
Rate of plate movement
rate = distance from ridge / age of that seafloor
Newly formed crust sits at the ridge (age 0). The farther the seafloor and the older it is, the faster the plate has spread. Convert units carefully — 1 km = 100,000 cm.
Texture, drainage, and retention
sand → high permeability, low water-holding · clay → low permeability, high water-holding · loam → balanced
Larger particles (sand) leave larger gaps, so water drains fast but little is retained. Smaller particles (clay) pack tightly, holding water but draining slowly.
Stream discharge
Q = A × v = (width × depth) × velocity
Q is discharge (volume per time, e.g. m³/s); A is the cross-sectional area of the channel; v is the flow velocity. More water, a deeper channel, or faster flow all raise discharge.
Normal Pacific vs. El Niño
normal: strong trade winds → upwelling off South America · El Niño: weak/reversed winds → upwelling shuts down
Follow the trade winds. Strong winds push warm water west and pull cold nutrient-rich water up in the east; weakened winds let warm water return east and suppress that upwelling.
Texture and behavior
sand → high permeability, low water and nutrient retention; clay → low permeability, high retention; loam → balanced
Permeability and retention trade off against each other, which is why loam is agriculturally best.
Rising and descending air
rising air cools → condenses → rain (equator, windward slopes); descending air warms → absorbs moisture → dry (30° latitude, leeward slopes)
One mechanism explains rainforests, deserts and rain shadows. Ask which way the air is moving.
Percent change in yield
percent change = [(new value − old value) / old value] × 100
A positive result is an increase, a negative result a decrease. Always divide by the ORIGINAL (old) value, not the new one.
Metal recovered from ore
mass of metal = mass of ore × (ore grade as a decimal)
Ore grade is the percent of the ore that is the target metal. Convert the percent to a decimal (0.5% = 0.005) before multiplying.
Maximum sustainable yield
sustainable harvest ≤ number of new individuals added per period
The maximum sustainable yield is the largest catch that can be taken repeatedly without shrinking the population. Harvest above the replacement rate and the stock declines toward collapse.
Ecological balance
footprint > biocapacity → ecological deficit (overshoot) · footprint < biocapacity → reserve
Compare demand (footprint) to supply (biocapacity). A footprint larger than the available biocapacity means resources are being drawn down faster than they regenerate.
Irrigation efficiency ranking
drip > spray/center-pivot > flood/furrow cost runs in the opposite order
Efficiency and salinization risk move together: the method that loses the least water also leaves the fewest salts.
Maximum sustainable yield
MSY occurs near half the carrying capacity, where population growth rate is greatest
Same logic as the logistic model in Unit 3. Harvest the surplus, not the stock.
Net energy (EROEI)
EROEI = energy returned / energy invested
The energy return on energy invested measures how much usable energy a source yields per unit of energy spent obtaining it. A higher EROEI means a better net-energy source; a ratio near 1 means the source is barely worth extracting.
Radioactive decay
amount remaining = initial amount × (1/2)ⁿ, where n = time / half-life
n is the number of half-lives that have elapsed. Each half-life halves the remaining amount: after n half-lives, the fraction left is (1/2)ⁿ.
Solar array power output
power output = area × solar intensity × efficiency
Area in m², solar intensity in W/m², efficiency as a decimal. The result is the electrical power in watts. Real panels convert only ~15–22% of incoming sunlight.
Energy use and cost
energy (kWh) = power (kW) × time (h); cost = energy (kWh) × price per kWh
Convert watts to kilowatts by dividing by 1,000 (1,500 W = 1.5 kW). Efficiency (%) = useful energy output / total energy input × 100.
Comparison criteria
emissions · other pollutants · land use · reliability · waste and decommissioning
Run all five on each source. A comparison that uses only emissions is incomplete by construction.
Efficiency through a chain
overall efficiency = product of the efficiencies of each step
0.35 generation × 0.95 transmission × 0.05 incandescent ≈ 1.7% of primary energy delivered as light.
Parts per million
1 ppm = 1 part per 1,000,000 = 0.0001% ; ppm = (parts of pollutant / total parts) × 1,000,000
ppm is a concentration ratio, not a mass. To convert ppm to a percent, divide by 10,000 (35 ppm ÷ 10,000 = 0.0035%).
Photochemical smog formation
NOₓ + VOCs + sunlight → ground-level ozone (O₃) + other secondary pollutants
The precursors (NOₓ and VOCs) come mostly from vehicle exhaust. Sunlight is essential, which is why photochemical smog peaks on hot, sunny afternoons.
pH is logarithmic
each 1-unit drop in pH = 10× more acidic (10× higher H⁺ concentration)
A difference of n pH units equals a factor of 10ⁿ in acidity. pH 3 vs pH 6 is a difference of 3 units = 10³ = 1,000 times more acidic.
Pollutant removal by a control device
mass removed = input mass × efficiency ; mass emitted = input mass × (1 − efficiency)
Express efficiency as a decimal (98% = 0.98). A 98%-efficient device still lets 2% of the pollutant escape.
Photochemical smog sequence
NO (morning peak) → NO₂ (midday peak) → O₃ (afternoon peak), driven by sunlight acting on NOₓ + VOCs
Ozone lags emissions by hours and peaks downwind, not at the source.
Pollutant concentration
concentration = mass of pollutant / volume of water
Keep units consistent. 1 g/L = 1,000 mg/L; concentrations in water are often reported in mg/L, which for dilute solutions is approximately equal to ppm.
The eutrophication cascade
excess N & P → algal bloom → algae die → decomposers ↑ → BOD ↑ → dissolved O₂ ↓ → fish die (dead zone)
The key insight: the harm is not the algae directly but the oxygen depletion caused by decomposers breaking down the dead algae. High BOD = low dissolved oxygen.
Waste generation and diversion
total waste = population × per-capita waste rate ; landfilled = total × (1 − recycling rate)
Multiply population by the per-person waste rate for the total, then subtract the recycled fraction to find what still reaches the landfill.
Biomagnification up the food chain
concentration at a level ≈ base concentration × (magnification factor)^(number of levels up)
If a toxin roughly multiplies by 10 at each trophic level, a predator three levels above the base carries about 10³ = 1,000 times the base concentration. LD50: lower value = more toxic.
Eutrophication chain
nutrients → algal bloom → light blocked → plants and algae die → decomposers consume O₂ → hypoxia and fish kill
The oxygen is consumed by decomposers, not by the algae. Omitting that step loses the causal point.
Catalytic ozone destruction
CFC + UV → free Cl· ; then Cl· destroys ozone repeatedly (one Cl → tens of thousands of O₃)
Chlorine is a catalyst: it is not used up, so a small amount of CFC destroys a vast amount of ozone. This is why even low CFC concentrations were so damaging.
CO₂-equivalent (using GWP)
CO₂-equivalent = mass of gas × GWP
GWP compares a gas to CO₂ (GWP = 1). Multiplying a gas’s mass by its GWP converts it to the mass of CO₂ that would trap the same heat.
Change over time (rate × time)
total change = rate of change × time
Useful for projecting impacts such as sea-level rise. If sea level rises at 3.3 mm per year, multiply by the number of years to estimate the total rise.
HIPPCO — causes of biodiversity loss
Habitat destruction · Invasive species · Population growth · Pollution · Climate change · Overexploitation
Habitat destruction is the #1 cause of extinction; invasive species are #2. Most threatened species suffer from several HIPPCO factors simultaneously.
Two distinct problems
stratospheric O₃ loss ← CFCs → more UV reaching the surface tropospheric GHGs → trapped infrared → warming
Different layer, different cause, different radiation, different consequence. CFCs happen to do both.
Ocean acidification chemistry
CO₂ + H₂O → H₂CO₃ → H⁺ + HCO₃⁻; the added H⁺ consumes CO₃²⁻, reducing carbonate for shell building
A chemistry consequence of CO₂, not a temperature one — which is why it would occur even without warming.

On the exam

How to get a 5

Key terms

Demographic Transition ModelStage 1: High birth/death rates. Stage 2: Death rates drop rapidly (medicine/sanitation), rapid growth. Stage 3: Birth rates drop. Stage 4: Low birth/death rates, stable population.
Tragedy of the commonsA shared, unregulated resource is overused because each user gains the whole benefit while sharing the cost — the model behind overfishing and air pollution.
Trophic Levels & 10% RuleOnly ~10% of energy is passed to the next trophic level; the rest is lost as heat (2nd Law of Thermodynamics).
Nitrogen Cycle: FixationConversion of atmospheric N2 into ammonia (NH3) or nitrates by soil bacteria (Rhizobium) or lightning, making it usable by plants.
EutrophicationNutrient enrichment causes an algal bloom; when the algae die, decomposers consume dissolved oxygen and fish suffocate, creating a hypoxic dead zone.
Ozone DepletionCFCs release chlorine atoms in the stratosphere. One Cl atom can destroy 100,000 O3 molecules, increasing UV radiation reaching Earth.
Photochemical smogNOx and volatile organic compounds react in sunlight to form tropospheric ozone. Worst on hot, sunny, windless days in traffic-heavy cities.
Greenhouse Gases (GHGs)CO2, CH4 (methane), N2O, CFCs, and water vapor. They trap outgoing infrared radiation, warming the troposphere.
Bioaccumulation vs biomagnificationBioaccumulation: a persistent, fat-soluble toxin builds up WITHIN one organism over its lifetime. Biomagnification: concentrations increase at each successive TROPHIC LEVEL, so top predators carry the highest burden. DDT, PCBs, and methylmercury are the standard examples.
K-selected vs r-selected speciesK-selected: few offspring, high parental care, late maturity (e.g., elephants). r-selected: many offspring, little care, early maturity (e.g., insects).
El Niño (ENSO)Weakening of trade winds in the Pacific. Warm water moves east toward South America, suppressing upwelling and altering global weather patterns.
Primary vs Secondary SuccessionPrimary: starts from bare rock (no soil), pioneer species like lichens. Secondary: starts after a disturbance (fire/clear-cutting) where soil is already present.
Simpson’s diversity indexD = 1 − Σ(n/N)², where n is the count of one species and N the total. Higher D means greater diversity. Σ(n/N)² alone is the dominance term — remember to subtract it from 1.
Rule of 70Doubling time (years) = 70 ÷ percent annual growth rate. Rate of natural increase = (crude birth rate − crude death rate)/10 expressed as a percent. A faster growth rate must always give a SHORTER doubling time.
Half-life calculationFraction remaining = (1/2)ⁿ, where n = elapsed time ÷ half-life. Count the number of half-lives first, then halve. Decay is asymptotic, so the quantity never reaches exactly zero.
Parts per million (ppm) and parts per billion (ppb)ppm = (mass of pollutant / total mass) × 10⁶; ppb uses 10⁹. Both masses must be in the same units first. In water, 1 mg/L is equivalent to 1 ppm because 1 L of water has a mass of about 1 kg.
EROEI (energy returned on energy invested)EROEI = energy returned ÷ energy invested. A ratio above 1 yields net energy; higher is better. Conventional oil and hydro rank high, while tar sands and corn ethanol rank low. Do not confuse EROEI with net energy, which is a difference rather than a ratio.
Global warming potential (GWP)The warming caused by one unit mass of a gas relative to CO₂ over a set horizon, usually 100 years. CO₂ = 1, methane ≈ 28, nitrous oxide ≈ 265, and some CFCs exceed 10,000. CO₂-equivalent = mass × GWP, so you MULTIPLY.
Removal efficiency of a treatment processEfficiency (%) = [(influent − effluent)/influent] × 100. The complementary quantity, effluent/influent, is the fraction remaining. A high-performing plant gives a high percentage removed and a low percentage remaining.
Stream dischargeQ = A × v = (width × average depth) × velocity, in m³/s. Check units: multiplying width by depth alone gives m², which cannot be a discharge. Discharge rises sharply after impervious-surface development because runoff reaches the channel faster.
Energy and cost calculations with kWhEnergy (kWh) = power (kW) × hours. Divide watt-hours by 1,000 to get kWh, then multiply by the rate in dollars per kWh. 1 kWh = 3.6 MJ, useful for converting between electricity and thermal energy units.
Power plant efficiency chainElectrical output = thermal input × efficiency, so thermal input = output ÷ efficiency. Fuel mass = thermal input ÷ energy density. Capacity factor = actual output ÷ nameplate output over a period; multiply nameplate capacity by capacity factor before using 8,760 hours per year.
Why the phosphorus cycle is differentPhosphorus has no significant gaseous phase, so it moves through rock weathering, soil, organisms, and sediments rather than the atmosphere. It cycles slowly and is usually the limiting nutrient in fresh water, which is why phosphate runoff triggers freshwater eutrophication.
Cultural eutrophication sequenceNutrient runoff (N and P) → algal bloom → algae die and sink → decomposer respiration consumes dissolved oxygen → hypoxia or a dead zone → fish kill. High BOD is the measurable signature of the oxygen-consuming organic load.